He was the subject of a BBC documentary last year, and was studied by naturalists throughout his life - including US expert Dian Fossey."
"He was born on 24 August 1974 and has been observed closely by researchers throughout his entire life," a statement from the Rwandan national parks office said.
"Tragically, he succumbed to old age on September 14."
The life expectancy of a healthy gorilla is about 40.
Tuesday, September 15, 2009
Titus "The Gorilla King" has Died
BBC News has reported that Rwanda is mourning its most famous mountain gorilla. The silverback Titus, known as 'The Gorilla King', has died at the age of 35.